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This list has some issues, Booking.com is Dutch and Philips' Consumer division is just a name that whitelabels other companies products and is owned by a Chinese investment company.
Booking.com is owned by an American company though
Also, Ecosia and Qwant rely on Bing for their search results I believe
or Google, but they are working together on their own search engine from what I heard
Booking is 100% owned by Priceline (renamed Booking Holdings). They only paid $113M for the company in 2005. That's a damn steal.
The other travel "alternatives" are either owned by Expedia, majority stock holders by Expedia, or just act as a front end search for Expedia and VRBO. It's honestly ridiculous.
Interesting, didn't know this. Thanks for pointing it out.